ok so, Pause is the land and sky wild life (any local bird species and local mammal), Etho is underwater (fish and one arctic fox because spirit reasons), and Beef is any domestic animal.
In last comic, Beef refers to time when humanity didn't exist (and he as well), meaning that they couldn't speak in words.
Yeah... that means Etho and Pause are kinda ancient. Btw they don't have these names, but I don't have any clue what to name those guys... It would probably be a nickname, or maybe i can just look in my local language vocabulary to merge some up xD